Large - scale hydrolysis of proteins occurs early in leaf senescence and is likely to dependent on the integrated action of groups of enzymes . the substrate for proteases in senescing leaves includes mainly enzymes and membrane proteins that function in photo synthesis 类囊体膜上结合的蛋白酶主要分为需能蛋白酶和不需能蛋白酶，参与类囊体膜蛋白和类囊体腔蛋白成熟时的加工以及外因导致蛋白质功能丧失时的降解。
