Studies by hargrave and geen estimated natural community graz - ing rates by measuring feeding rates of individual zooplankton species in the laboratory and then computing community grazing rates for field conditions using the known population density of grazers 由哈格雷夫( hargrave )和吉恩( geen )所进行的研究，对自然条件下的群落食草比例进行了估计，其手段是通过测量出实验室内单独的浮游动物种类的结食比例，然后利用已知的食草动物种群密度，计算出实地状况下的群落食草比例。
